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Freeing the hens: Workshop outcomes for applying
ethology to the development of cage-free
housing systems in the commercial egg industry
T.B. Rodenburg, M.F. Giersberg,
P. Petersan, S. Shields
Pampered pets or poor bastards? The welfare of
dogs kept as companion animals
I. Meyer, B. Forkman,
M. Fredholm, C. Glanville,
B. Guldbrandtsen, E. Ruiz Izaguirre,
C. Palmer, P. Sandøe
Timing of part-time group housing for
farm rabbits: effects on reproductive
performance, skin injuries and behaviour
L.G.W. Van Damme, E. Delezie,
B. Ampe, F.A.M. Tuyttens
Associations between osteoarthritis
and duration and quality of
night-time rest in dogs
M. Smith, M. Mendl,
J.C. Murrell
Tracking performance in poultry is affected by data
cleaning method and housing system
L. Candelotto, K.J. Grethen,
C.M. Montalcini, M.J. Toscano,
Y. Gómez- Effect of cooperative care training on physiological
parameters and compliance in dogs
undergoing a veterinary examination
L. Wess, A. Böhm,
M. Schützinger, S. Riemer,
J.R. Yee, N. Affenzeller,
C. Arhant Responsiveness of domesticated goats towards
various stressors following long-term
cognitive test exposure
K. Rosenberger, M. Simmler,
J. Langbein, C. Nawroth,
N. Keil
Predictors for plumage damage and bloody
lesions indicative of feather pecking
in pullets reared in aviaries
C. Mels, K. Niebuhr,
A. Futschik, J.-L. Rault,
S. Waiblinger
Does the duration of repeated temporary separation
affect welfare in dairy cow-calf
contact systems?
N. Roadknight, W. Wales,
E. Jongman, P. Mansell,
G. Hepworth, A. Fisher- Bark to the Future: The welfare of domestic dogs
during interaction with a positively
reinforcing artificial agent
N. Shaw, F. Wemelsfelder,
L.M. Riley - Consumer attitudes towards farm animal welfare
in Argentina, Chile, Colombia, Ecuador,
Peru and Bolivia: A segmentation-based study
L.X. Estévez-Moreno, G.C. Miranda-de la Lama,
G.G. Miguel-Pacheco 
Commercial broiler breeder pullet hens use
perches but show no preference for
perch type or height
G. Vasdal, S.G. Gebhardt-Henrich,
K.E. Kittelsen, F.M. Tahamtani
Conventional laboratory housing increases morbidity
and mortality in research rodents:
results of a meta-analysis
J. Cait, A. Cait,
R.W. Scott, C.B. Winder,
G.J. Mason- Commercial Free-Range Laying Hens’ Preferences for Shelters
with Different Sunlight Filtering Percentages
S. Rana, C. Lee,
J.M. Lea, D.L.M. Campbell - Application of Ultraviolet Light for Poultry Production:
A Review of Impacts on Behavior,
Physiology, and Production
S. Rana, D.L.M. Campbell 
Music modulates emotional responses in growing pigs
J. Zapata Cardona, M.C. Ceballos,
A.M. Tarazona Morales, E. David Jaramillo,
B.D.R. Rodríguez
Canine-supported bio detection of corpses, anatomical sections
and biological fluids at crime scenes
M.A. Romero Luna, G.A. Bohórquez,
J.U. Rojas-Guevara, J.A. Baracaldo Eusse,
D. Alit Mansegosa
Insect-repelling behaviour in horses in relation to insect
prevalence and access to shelters
J.W. Christensen, C. G. Strøm,
K. Nicová, C.L. deGaillard,
P. Sandøe, H. Skovgårdg- Relationships between owner and household characteristics
and enrichment and cat behaviour
I. Windschnurer, A. Häusler,
S. Waiblinger, G.J. Coleman 
Environmental Enrichment for Rats and Mice Housed
in Laboratories: A Metareview
A.S. Ratuski, D.M. Weary
- Qualitative behaviour assessment as part of a welfare assessment in
flocks of laying hens
G. Vasdal, K. Muri,
S.M. Stubsjøen, R.O. Moe,
K. Kittelsen 
Lying in spatial proximity and active social behaviours capture
different information when analysed
at group level in indoor-housed pigs
I. Camerlink, K. Scheck,
T. Cadman, J.-L. Rault
Goats from different selection lines
differ in their behavioural flexibility
C. Nawroth, K. Rosenberger,
N. Keil, J. Langbein
A method for assessing sustainability,
with beef production as an example
D.M. Broom
Modulating captive mammalian social behavior:
A scoping review on olfactory treatments
A.J. Barabas, S.R. Dijak,
J.F. Yatcilla, D.N. Walker,
B.N. Gaskill
Once bitten, twice shy: aggressive and defeated
pigs begin agonistic encounters
with more negative emotions
L. Oldham, G. Arnott,
I. Camerlink, A. Doeschl-Wilson,
M. Farish, F. Wemelsfelder,
S.P. Turner
Hucul horses’ learning abilities in different learning
tests and the association with behaviour,
food motivation and fearfulness
K. Olczak, C. Klocek,
J.W. Christensen
Horses’ (Equus caballus) Ability to Solve Visible but Not Invisible
Displacement Tasks Is Associated With
Frustration Behavior and Heart Rate
M.V. Rørvang, K. Nicova,
H. Sassner, C. Nawroth
- Live black soldier fly larvae provisioning
is a promising environmental enrichment
for pigs as indicated by feed- and
enrichment-preference tests
A.F. Ipema, W.J. Gerrits,
E.A. Bokkers, B. Kemp,
J.E. Bolhuis 
Measuring Calves’ Usage of Multiple Environmental
Enrichment Objects Provided Simultaneously
A.C. Strappini, G. Monti,
P. Sepúlveda-Varas, I. de Freslon,
J.M. Peralta
Performance in cognitive tasks is not affected
by long-term cognitive test exposure
in conceptually different tests
K. Rosenberger, M. Simmler,
J. Langbein, N. Keil,
C. Nawroth
Learned control of urinary reflexes
in cattle to help reduce greenhouse gas emissions
N. Dirksen, J. Langbein, J.,
L. Schrader, B. Puppe,
D. Elliffe, K. Siebert,
V. Röttgen, L. Matthews
- Calf-directed affiliative behaviour
of dairy cows in two types of
cow-calf contact systems
M.L. Wenker, C.G. van Reenen,
D. de Oliveira, K. McCrea,
C.M. Verwer, E.A. Bokkers - How would dogs spend their days?
Activity patterns in four populations
of domestic dogs
S. Griss, S. Riemer,
C. Warembourg, et al. 
A rein tension signal can be reduced
by half in a single training session
M. Eisersiö, J. Yngvesson, A. Byström,
P. Baragli, A. Egenvall
Impact of digital dermatitis on
feedlot cattle behaviour
A.D. Thomas, K. Orsel,
J.A. Cortés, E.A. Pajor
- Compounds from plantar foot sweat, nesting material,
and urine show strain patterns associated
with agonistic and affiliative behaviors
in group housed male mice, Mus musculus
A.J. Barabas, H.A. Soini,
M.V. Novotny, D.R. Williams,
J.A. Desmond, J.R. Lucas,
M.A. Erasmus, H.W. Cheng,
B.N. Gaskill 
Association patterns across multiple gestation
cycles within a dynamic sow pen
K. Horback, C. McVey,
M. Pierdon
The role of canine human remains detection"
teams in criminal investigation
J.U. Rojas-Guevara, J.D. Córdoba-Parra,
G.A. Bohórquez, R.A. Vega-Contreras,
P. Prada-TiedemannEffects of restraint on heifers during
gentle human-animal interactions
A. Lange, S. Waiblinger,
R. van Hasselt, R. Mundry,
A. Futschik, S. Lürzel
- Tail-biting behaviour pre-weaning:
Association between other pig-directed
and general behaviour in piglets
F. Hakansson, J.E. Bolhuis - Rooting for feed: Mixing corn pellets into rooting
material increases the use but not cleanliness
of rooting areas for growing-finishing pigs
M. Knoll, E.A.M. Bokkers,
C. Leeb, C. Wimmler,
H. Mai-Lis Andersen, R. Thomsen,
B. Früh, M. Holinger 
The potential of feeding patterns to assess generic
welfare in growing-finishing pigs
J.D. Bus, I.J.M.M. Boumans,
L.E. Webb, E.A.M. Bokkers- Who's the Boss? Assessing Convergent Validity of
Aggression Based Dominance Measures in
Male Laboratory Mice, Mus Musculus
A.J. Barabas, J.R. Lucas,
M.A. Erasmus, H.W. Cheng,
B.N. Gaskill
Maternal deprivation affects
goat kids’ stress coping behaviour
C. Toinon, S. Waiblinger,
J.-L. Rault- Chewing behaviour in dogs –
a survey-based exploratory study
C. Arhant, R. Winkelmann,
J. Troxler - Providing environmental enrichments affects activity
and performance, but not leg health
in fast- and slower-growing broiler chickens
I.C. de Jong, X.E. Blaauw,
J.A.J. van der Eijk, C.S. da Silva,
M.M. van Krimpen, R. Molenaar,
H. van den Brand 
Validation of a real-time location system
for zone assignment and neighbor
detection in dairy cow groups
N. Melzer, B. Foris,
J. Langbein
Milk- and starter-feeding strategies
to reduce cross sucking in pair-housed
calves in outdoor hutches
R.S. Salter, K.J. Reuscher,
J.M.C. Van Os
The Legislative, Ethical, and Conceptual
Importance of Replicability in
Farm Animal Welfare Science
C. Nawroth, L. Gygax- Are free-ranging Kune Kune pigs
(Sus scrofa domesticus) able to solve
a cooperative task?
M. Koglmüller, M. Wondrak,
I. Camerlink, J.-L. Rault,
L. Huber - Preference testing for UV light
spectrum and intensity in laying hens
M.S. Rana, A.M. Cohen-Barnhouse,
C. Lee, D.L. Campbell
Does coping style affect behavioral
responses and growth performance
of lambs weaned at different ages?
C. Çakmakçı, S. Karaca,
G.A. María- Decomposition Residual Odor Volatiles in
Soil from a West Texas Environment
J. Raymer, P.A. Prada-Tiedemann,
J.U. Rojas-Guevara 
Individual Detection and Tracking
of Group Housed Pigs in Their Home Pen
Using Computer Vision
L.E. Van Der Zande, O. Guzhva,
T.B. Rodenburg- Effects of hatching on-farm on behaviour,
first week performance, fear level
and range use of organic broilers
C.T. Jessen, L. Foldager,
A.B. Riber
Happiness, Welfare, and Personality
in Rhesus Macaques (Macaca mulatta)
L.M. Robinson, N.K. Waran,
I. Handel, M.C. Leach- Piling behaviour in Swiss layer flocks:
description and related factors
J. Winter, M. Toscano,
A. Stratmann - Vocalisations in Farm Animals:
a Step Towards Positive Welfare Assessment
K.A. Laurijs, E.F. Briefer,
I. Reimert, L.E. Webb - Affective styles and emotional
lateralization: A promising framework
for animal welfare research
C. Goursot, S. Düpjan,
B. Puppe, L.M. Leliveld
- Caretaker attitudes and animal training are
associated with alpaca behaviour towards humans—
An online survey
I. Windschnurer, L. Fischer,
T. Yanagida, C. Eibl,
S. Franz, S. Waiblinger 
Influence of Different LED Light Colour
Temperatures on the Preference Behaviour
of Weaned Piglets
S. Goetz, C.M.C. Raoult,
K. Reiter, M. Wensch-Dorendorf,
D. Werner, E. v. Borell
Detection of cow hind-leg activity
during milking by using a 3-dimensional
accelerometer attached to the milking cluster
C.M.C. Raoult, A.A. Margerit,
S. Fricker, E.F. Bluemel,
P.E. Savary
Goats show higher behavioural flexibility
than sheep in a spatial detour task
C.M.C. Raoult, B. Osthaus,
A.C.G. Hildbrand, A.G. McElligott,
C. Nawroth
- Rider effects on horses' conflict behaviour,
rein tension, physiological measures
and rideability scores
J. Winther Christensen, R. Munk,
L. Hawson, R. Palme, T. Larsen,
A. Egenvall, U. U. König von Borstel,
M. Vilain Rørvang 
Access to chewable materials
during lactation affects sow behaviour
and interaction with piglets
K.-M. Swan, H. Telkänranta,
C. Munsterhjelm, O. Peltoniemi,
A. Valros
Development of existing Scoring Systems
to Assess Behavioural Coping
in Shelter Cats
E. N. Hirsch, M. Andersson,
J. Loberg, L. M. Lidfors
Familiarity influences social networks
in dairy cows after regrouping
B. Foris, H.-G. Haas,
J. Langbein, N. Melzer
- Balancing skill against difficulty -
behavior, heart rate and heart rate variability
of shelter dogs during two different
introductions of an interactive game
C. Arhant, B. Altrichter, S. Lehenbauer,
S. Waiblinger, C. Schmied-Wagner, J. Yee 
Gentle interactions with restrained
and free-moving cows: Effects on
the improvement of the
animal-human relationship
A. Lange, S. Waiblinger,
A. Heinke, K. Barth,
A. Futschik, S. Lürzel- How Can Cattle Be Toilet Trained?
Incorporating Reflexive Behaviours
into a Behavioural Chain
N. Dirksen, J. Langbein,
L. Schrader, B. Puppe,
D. Elliffe, K. Siebert,
V. Roettgen, L. Matthews
contrafreeloading task
K. Rosenberger, M. Simmler,
C. Nawroth, J. Langbein,
N. Keil
- Rewarding memories? Behaviour of broiler
chickens towards peat
in flocks with and without
previous exposure to peat
J. Vas, N. BenSassi,
G. Vasdal, R. C. Newberry
Reactions to Different Auditory
Stimuli During Gentle
Human-Animal Interactions
A. Lange, L. Bauer,
A. Futschik, S. Waiblinger,
S. Lürzel
The effect of perches and
aviary tiers on the mating behaviour
of two hybrids of broiler breeders
S. G. Gebhardt-Henrich, A. Jordan,
M. J. Toscano, H. Würbel
Early feeding experiences of
piglets and their impact on novel
environment behaviour and
food neophobia
A. Middelkoop, B. Kemp, E. Bolhuis
Understanding allogrooming through
a dynamic social network approach:
An example in a group
of dairy cows
I. de Freslon, J.M. Peralta
A.C. Strappini, G. Monti
Effect of cow-calf contact
on cow motivation to reunite
with their calf
M.L. Wenker, et al.
Sensory abilities of horses
and their importance to
equitation science
M.V. Rørvang, B.L. Nielsen,
A.N. McLean- Comparative assessment of
general behaviour and fear-related
responses in hatchery-hatched and
on-farm hatched broiler chickens
M.F. Giersberg, et al.
Short- and long-term
effects of rearing dairy calves
with contact to their mother
on their reactions toward humans
S. Waiblinger, et al.
Play and social behaviour
of calves with or without
access to their dam
and other cows
S. Waiblinger, et al.- Intra-group lethal gang
aggression in domestic pigs
(Sus scrofa domesticus)
I. Camerlink, J.-Y. Chou,
S.P. Turner - Emotional contagion and
its implications for animal welfare
S. Düpjan, A. Krause,
L.R. Moscovice, C. Nawroth
- Provisioning of live
black soldier fly larvae
(Hermetia illucens)
benefits broiler activity
and leg health in a
frequency- and dose-
dependent manner
A.F. Ipema, et al. - Perch positioning affects
both laying hen locomotion
and forces experience
at the keel
C. Rufener, A.K. Rentsch,
A. Stratmann, M.J. Toscano - Pullet rearing affects
collisions and perch use
in enriched colony cage
layer housing
A.N. Pullin, et al. - Current state of
knowledge on the cognitive
capacities of goats and its
potential to inform species-
specific enrichment
G. Zobel & C. Nawroth
